Elevation to species level of Nebria (Oreonebria) tresignore (Szallies & Huber, 2014) stat. nov. (Coleoptera: Carabidae), a cold-adapted, endemic and threatened ground beetle of the Italian Alps

Nebria (Oreonebria) tresignore (Szallies & Huber, 2014) stat. nov. a ground beetle living in high-altitude habitats of a restricted area of the Italian Alps (i.e., Orobic Alps), previously considered a subspecies of N. (Oreonebria) soror K. Daniel, 1903, was phylogenetically analyzed based on the combination of nuclear (Tpl and WG) and mitochondrial (COIa, COIb and 16S) markers. Given the morphological (male genitalia), biogeographical and ecological differences with closely related species and the newly performed phylogenetic reconstruction, its elevation to species level is proposed. This cold-adapted species is at high risk of extinction due the ongoing climate warming and glaciers retreat on the Italian Alps.